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I I \ ,_Grazers \ \ 5 \ 0/,°\ Deposit Feeders . I ... ~~~ . 001 -=;; . 0 Individual Weight Fig. 10. Relation between macrofaunal biomass (g m - 2) and individual weight (g in ashfree dry weight) in June 1980 on a sandy flat in Konigshafen, island of Sylt. Mixedfeeders refers to a combination of suspension- and deposit-feeding. (After H. 6 Production and Energy Flow H. Asmus and R. Asmus (1985) measured primary and secondary production on three tidal flats in Konigshafen. 4). In areas where the annual Zostera noltii forms dense patches during summer, gross primary productivity is much higher than in the flats with micro algae only.

However, in many years the small but very numerous diatom-grazing snail Hydrobia ulvae dominates the biomass with about 10 gem - 2 (H. Asmus and R. Asmus 1985; Fig. 10). 3. Abundance (individuals below 1 m 2 ) of macrofauna at 4 sites in April 1975, Konigshafen, island of Sylt. 0mm High flat 30 Grass bed 50 Sand flat 70 Mud flat 80 Anthozoa Sagartiogeton viduata (0. F. ) Nereis diversicolor O. F. ) Seoloplos armiger (0. F. Miiller) Magelona mirabilis (Jones 1977) Spio filicornis (0. F. ) Tharyx marioni (Saint-Joseph) Ampharete acutifrons (Grube) Lanice conchilega (pallas) Fabricia sabella Ehrenberg Oligochaeta Tubificoides benedeni (D'Udekem) Bivalvia Mytilus edulis L.

Common life cycle in epibenthic decapod crustaceans and demersal fish living in coastal regions. Pelagic larvae develop offshore, drift inshore and metamorphose to benthic juveniles on the landward flats. Early stages bury at low tides while later stages migrate back and forth with the tides. Adults forage in the subtidal. The tidal flats function as nursery grounds Low tide preference is not only shown by organisms of terrestrial origin. Some of the micro algae living on tidal flats exhibit vertical migrations within the sediment.

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